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Computational modeling of the macaque brain grounds hypotheses on the brain of LCA-m (the last common ancestor of monkey and human). Elaborations thereof provide a brain model for LCA-c (c for chimpanzee). The Mirror System Hypothesis charts further steps via imitation and pantomime to protosign and protolanguage on the path to a "language-ready brain" in Homo sapiens, with the path to speech being indirect. The material poses new challenges for both experimentation and modeling.
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